Bu bölgede aktivatörler ve represörler etkinlik gösterir. Represörler bu bölgeye bağlanarak RNA Polimerazın bağlanmasını engellerler ve böylece transkripsiyona mani olurlar. Bu bölge promoter bölgenin sağında bulunur. (Resimde "O" harfi ile gösterilen bölge)
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